This comparison can be between two different habitats or a comparison of one habitat over time. Application of shannons index to study diversity with reference to census data of assam. Shannon wiener index of diversity information index a measure used by ecologists when a system contains too many individuals for each to be identified and examined. For categorical data, there is no mean or median and so the measures of variation described in measures of variability cant be used. Shannonwiener index h most commonly used index of diversity in ecological studies. If i obtain the shannonweiner diversity index as 2. This index incorporates both species richness and species evenness. The shannon wiener index is defined as is the natural logarithm of the total number of species. Lets use r to calculate h for the two communities in the example above.
The maximum diversity hmax of a sample is found when all species are equally abundant. This calculator is free to use and is designed for biologists, ecologists, teachers, and students needing to quickly calculate the biodiversity indexes of an ecosystem. Diversity indices like the shannon entropy shannon wiener index and the ginisimpson index are not themselves diversities. Species richness, diversity and abundance of some decapod crustaceans in coastal waters of ondo state, south west, nigeria.
According to local occurrence status, there were 12 species ranked as very abundant, 19 abundant, 62 very common, 16 common and 41 fairly common. Hence, for a fixed simpson diversity index, d, the shannon wiener diversity index can take on at least all values greater than or equal to log1d. Wikipedia the degree of variation of life forms within a given species, ecosystem, biome, or an entire planet. Chesapeake bay benthic index of biotic integrity tidal monitoring and analysis workgroup june 12, 2012. While it is fully termed the shannonwiener function, after being derived independently by claude shannon and norbert wiener 10,11, it is often called the shannonweaver index after wallace weaver, shannons coauthor. It uses the logarithm, and we will write it here with the natural logarithm. A widely used dominance index is simpsons diversity index. Comparing species diversity and evenness indices by c. Pdf prediction of the shannon wiener diversity index. This makes the shannon index and the simpson diversity or gini coefficient special cases of a more general index. A measure used by ecologists when a system contains too many individuals for each to be identified and examined. Sample values s 60,10,25,1,4 number of species n 5 first, let us calculate the sum of the given values. Shannon wiener species diversity index formula data analysis.
Where p i is the relative abundance of species i, s is the total number of species present and ln is the natural log. The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field. The intuitive significance of this index can be described as follows. There were, however, 7 species, in each category, recorded as uncommon. Simpson diversity and the shannon wiener index as special cases of a generalized entropy. Online biodiversity calculator which helps to calculates the shannon s diversity index and evenness from the given sample data. In particular, for a random sample, we can use shannons index of diversity aka as shannonweiners index, which is.
Ali faculty of veterinary sciences, bahauddin zakariya university multan, pakistan. In the following we will use the shannon diversity index h in other applications also named shannon entropy which is based on the geometric mean, and the true diversity of order one. The shannon index is a measurement used to compare diversity between habitat samples. This index is borrowed from information science, and is calculated as follows. On some occasions it is called the shannon wiener index and on other occasions it is called the shannon weaver index. A tribute to claude shannon 19162001 and a plea for. Application of shannons index to study diversity with.
Using ecological diversity measures with bacterial communities. Congress office of technology assessment the variety and variability. Can i get a single index value for each cave even if the methods are different. Benthic index of biotic integrity metric scoring system. Aug 28, 2006 also, that shannon wiener diversity approaches log1d as p approaches 0. Shannon wiener index biodiversity studies pdf species diversity of ecological communities is complicated. Shannon wiener index is the most widely index which used in marine biodiversity. And what about that caves which have stream or pools where the fauna can be very abundant. Diversity indices that combine both richness and abundance i. Prediction of the shannon wiener diversity index from trace element profiles in sediments around the statfjord platforms.
Diversity of classes shannon, simpson, species statsdirect. This tutorial explains how to calculate the shannon wiener diversity index and evenness. Pages 334338, in tested studies for laboratory teaching, volume 27 m. Shannons diversity index real statistics using excel. Also, that shannon wiener diversity approaches log1d as p approaches 0. A diversity index is a mathematical measure of species diversity in a community. First, enter the number of species, and then enter the name you wish to give the species, if available, and the given populations for each of the speciesin any given order. There seems to be some confusion in the literature and both are commonly used for the same index. This is by far the most widely used diversity index. Biodiversity merriamwebster the existence of many different kinds of plants and animals in an environment. This diversity measure is based on information theory. Species richness, diversity and abundance of some decapod.
View enhanced pdf access article on wiley online library html view download pdf for offline viewing. Brief outlines of the two types of diversity indices of biodiversity are discussed in this article. Diversity indices provide more information about community composition than simply species richness i. The shannon weaver model the shannon weaver model is typical of what are often referred to as transmission models of communication. Biodiversity calculator this calculator is free to use and is designed for biologists, ecologists, teachers, and students needing to quickly calculate the biodiversity indexes of an ecosystem. Calculate the shannon diversity index and evenness for these sample values. Sample values s 60,10,25,1,4 number of species n 5. Instead we use a measure of the distribution of observations among the categories.
How to calculate shannon wiener diversity index definition. The actual formula for the shannonweiner diversity index is. One index value on its own holds very little value. Shannonwiener index of diversity information index source. Apr 10, 2020 the shannonweaver is the preferred term. C the more species you have, the more diverse the area. Interestingly shannon, a physicist, developed the index as a formula for measuring the entropy of matter in the universe. A a diversity index is a mathematical measure of species diversity in a given community. The shannon index has been a popular diversity index in the ecological literature, where it is also known as shannons diversity index, the shannonwiener index, the shannonweaver index and the shannon entropy.
Introduction the shannon index has been a popular diversity index in the ecological. We suggest an explanation for this dual use of terms and in so doing we offer a tribute to the late claude shannon who passed away on 24 february 2001. Historically, the index has been used to measure the effects of habitat quality such as effects of pollution effluents. It combines both evenness and richness in a single measure. Second, they calculated the shannon wiener index of biodiversity for each park. Demonstration of how to calculate the shannonwiener diversity index in microsoft excel. The actual formula for the shannon weiner diversity index is. Simpson index must also be calculated for this will give you a general idea regarding dominance. First, enter the number of species, and then enter the name you wish to give the species, if available, and the given populations for each of the species. The most common measures of biodiversity are species richness, simpsons index and shannon wiener index. Habitat class metric tf ol lm hm sand hm mud po sand po mud shannon wiener species diversity index x x x x x total species abundance x x x x x x x total species biomass. Dpsi log pi, where s is the total number of species in the sample, i is the total number of individuals in. Biodiversity calculator for the simpson and shannon indexes. It turns out that the mathematical relationships hold true whether one is dealing with molecules in solution or.
In fact, in the late 1940s shannon had built on the work of wiener. Measuring diversity has been of historical significance and due to the obvious declines in habitat diversity this data is still valuable. Join facebook to connect with shannon wiener and others you may know. To do this, the proportion of each species of the total is calculated, the. That value of log1d looks like a hard minimum, but proof eludes me this morning. On some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index. Statistics shannon wiener diversity index tutorialspoint. For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot. What is the correct name for the biodiversity index. Is there other information that could be used to find the correct name for the index, if one exists.
First, let us calculate the sum of the given values. The shannon index increases as both the richness and the evenness of the community increase. Natural resource biometrics measures of diversity shannon. Biased richness and evenness relationships within shannon. The uncertainty surrounding the shannon index is exemplified by the variety of names given to it. Dominance indices are weighted toward the abundance of the commonest species. P i fraction of the entire population madeup of species i proportion of a species i relative to total number of. To measure the tree species diversity of a forest you need to know the total area of your forest and the area occupied by each species.
I found that the diversity index of zooplankton from a recent investigation to be 1. Thirtytwo richnessbased thirdorder polynomial regression models r 0. Shannon wiener species diversity index calculation. Shannon index is a commonly used diversity index that takes into account. H has no intuitive interpretation in terms of probability and is sensitive to sample size. In particular, for a random sample, we can use shannon s index of diversity aka as shannon weiner s index, which is defined as where n i is the number of observations from the sample in the i th of k nonempty categories and n is the sample size.
A tribute to claude shannon 19162001 and a plea for more. I was thinking to use the shannon wiener diversity index and to use both pitfall traps and timed area search sampling methods. Lets compute the shannonweiner diversity index for the same hypothetical community in the previous example. The purpose of this analysis was to empirically model and graphically illustrate the numerical relationships between richness s, 435 species and evenness e with respect to shannonwiener index h. This is the simplest of all the measures of species diversity. Posted on 201021 20170904 author bpmsg categories online tools, other articles tags berger parker, calculation, excel, shannon entropy, shannon weaver index, shannonwiener index, simpson index diversity as business kpi alpha and beta diversity. Regarding your result, a shannon wiener diversity index of 2. A greater number of species and a more even distribution both increase diversity as measured by h. Shannon s diversity index shannon s diversity index is simply the ecologists name for the communication entropy introduced by claude shannon. What is the definition of shannonwiener diversity index.
Statistics shannon wiener diversity index in the literature, the terms species richness and species diversity are sometimes used interchangeably. We suggest that at the very least, authors should define. Simpson diversity and the shannonwiener index as special. The shannonwiener diversity index is one measure that we will use to try to. The pros and cons of the shannon and weaver model free essays. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. B based on the species richness the number of species present and species abundance the number of individuals per species.
Engels department of zoology, state university of ghent, belgium in a low diversity brackish water habitat the diversity of the copepod community is best measured with the shannon wiener information function and its evenness by an index proposed by the first author eh181. An example of the shannon wiener diversity index for science students. Jan 26, 2012 shannon index is a commonly used diversity index that takes into account both abundance and evenness of species present in the community. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. The purpose of this analysis was to empirically model and graphically illustrate the numerical relationships between richness s, 435 species and evenness e with respect to shannon wiener index h. The fact that the index incorporates both components of biodiversity can be seen as both a strength and a weakness. The shannonweiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. A value for evenness approaching zero reflects large differences in abundance of species, whereas an evenness of one means all species are equally abundant.
Key wordsdiversity indices, shannon index, shannon wiener index, shannonweaver index, species richness, species diversity. Department of zoology, university of the punjab, lahore, pakistan corresponding author email. Shannon s index of diversity h is derived from information theory, originally in the context of information in telephone systems shannon, 1948. They are just indices of diversity, in the same way that the diameter of a sphere is an index of its volume but is not itself the volume.